Effects of hippocampal manipulations on the classically conditioned nictitating membrane response: simulations by an attentional-associative model. 1989

N A Schmajuk, and J W Moore
Department of Psychology, Northwestern University, Evanston, IL 60201.

The effects of various hippocampal manipulations on the classically conditioned nictitating membrane (NM) response were simulated by a real-time attentional-associative model. The model incorporates: (a) a mechanism capable of establishing associations between conditioned (CS) and unconditioned stimuli (US) and between two CSs, (b) a mechanism that combines CS-CS and CS-US associations and builds 'computational cognitive maps', (c) an attentional rule that 'tunes in' relevant CSs and 'tunes out' irrelevant CSs, (d) performance rules that convert learning variables into topography of the rabbit NM response, and (e) rules that convert learning variables into neuronal firing. The present study explored three hypotheses regarding hippocampal function: (a) hippocampal lesions (HL) impair the 'tuning out' of irrelevant stimuli, (b) hippocampal stimulation (HS) and hippocampal long-term potentiation (LTP) facilitate the 'tuning in' of relevant stimuli, and (c) hippocampal unit activity is proportional to the instantaneous value of associative variables that participate in the 'tuning' mechanisms. Computer simulations for the HL case were carried out for conditioning under different interstimulus intervals (ISI) with different types of US, discrimination reversal, and sensory preconditioning. Computer simulations for the HS and LTP case were carried out for acquisition of delay conditioning and of classical discrimination. In addition, simulations of hippocampal unit activity during acquisition and extinction are presented. Under the 'tuning out' hypothesis for the HL case, the model proved capable of simulating a large portion of the experimental data, showing discrepancies with relevant literature only when describing HL effects on trace conditioning with shock US under short and long ISIs, trace conditioning with air puff US under long ISIs, discrimination reversal and sensory preconditioning. Under the 'tuning in' hypothesis regarding the HS and LTP case, the model proved capable of simulating acquisition of delay conditioning but not acquisition of classical discrimination. Under the assumption that neuronal activity is proportional to the instantaneous value of associative variables, the model is able to simulate hippocampal unit activity.

UI MeSH Term Description Entries
D007858 Learning Relatively permanent change in behavior that is the result of past experience or practice. The concept includes the acquisition of knowledge. Phenomenography
D008568 Memory Complex mental function having four distinct phases: (1) memorizing or learning, (2) retention, (3) recall, and (4) recognition. Clinically, it is usually subdivided into immediate, recent, and remote memory.
D008570 Memory, Short-Term Remembrance of information for a few seconds to hours. Immediate Recall,Memory, Immediate,Working Memory,Memory, Shortterm,Immediate Memories,Immediate Memory,Immediate Recalls,Memories, Immediate,Memories, Short-Term,Memories, Shortterm,Memory, Short Term,Recall, Immediate,Recalls, Immediate,Short-Term Memories,Short-Term Memory,Shortterm Memories,Shortterm Memory,Working Memories
D008959 Models, Neurological Theoretical representations that simulate the behavior or activity of the neurological system, processes or phenomena; includes the use of mathematical equations, computers, and other electronic equipment. Neurologic Models,Model, Neurological,Neurologic Model,Neurological Model,Neurological Models,Model, Neurologic,Models, Neurologic
D009433 Neural Inhibition The function of opposing or restraining the excitation of neurons or their target excitable cells. Inhibition, Neural
D009434 Neural Pathways Neural tracts connecting one part of the nervous system with another. Neural Interconnections,Interconnection, Neural,Interconnections, Neural,Neural Interconnection,Neural Pathway,Pathway, Neural,Pathways, Neural
D009474 Neurons The basic cellular units of nervous tissue. Each neuron consists of a body, an axon, and dendrites. Their purpose is to receive, conduct, and transmit impulses in the NERVOUS SYSTEM. Nerve Cells,Cell, Nerve,Cells, Nerve,Nerve Cell,Neuron
D011817 Rabbits A burrowing plant-eating mammal with hind limbs that are longer than its fore limbs. It belongs to the family Leporidae of the order Lagomorpha, and in contrast to hares, possesses 22 instead of 24 pairs of chromosomes. Belgian Hare,New Zealand Rabbit,New Zealand Rabbits,New Zealand White Rabbit,Rabbit,Rabbit, Domestic,Chinchilla Rabbits,NZW Rabbits,New Zealand White Rabbits,Oryctolagus cuniculus,Chinchilla Rabbit,Domestic Rabbit,Domestic Rabbits,Hare, Belgian,NZW Rabbit,Rabbit, Chinchilla,Rabbit, NZW,Rabbit, New Zealand,Rabbits, Chinchilla,Rabbits, Domestic,Rabbits, NZW,Rabbits, New Zealand,Zealand Rabbit, New,Zealand Rabbits, New,cuniculus, Oryctolagus
D011939 Mental Recall The process whereby a representation of past experience is elicited. Recall, Mental
D003214 Conditioning, Classical Learning that takes place when a conditioned stimulus is paired with an unconditioned stimulus. Reflex, Conditioned,Classical Conditioning,Classical Conditionings,Conditioned Reflex,Conditionings, Classical

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